Cylinders go lean for a few reasons, lack of fuel (Bad injector, low pressure, low electrical command either from programming or from an issue such as high resistance.) or an air imbalance. Depending on how the air box is designed one cylinder could be getting far more air than the others. Intake port design, injector angle, etc all play a role as well but at the 374HP level odds are that's still stock, maybe ported. Something else, during the reassembly did they check the valve springs? If you have a lazy intake valve spring on 4 under boost the valve could be getting held open causing more air to rush in before it closes. Basically the valve doesn't follow the cam lobe, the boost pressure holds it open longer forcing more air in, once the piston starts coming up the pressure balances and the valve closes. Are you running stock springs? If so that's exactly where I'm going to look. Factory springs have enough open pressure to handle the valve's weight but not the valves weight with 17PSI pushing on it.
On to the next round of questions, along the way did you do plug checks? How do the plug checks look? If everything's balanced then all the plugs will look the same. Lets start there and see what you've got going on.